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Moreover, the expression of key spermatogenesis markers, including UCHL1, PLZF, SCP3, and PCNA, which were reduced in aged testes, were significantly restored or increased following FOXO4-DRI treatment

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doi: 10.3389/fimmu.2026.1779543 Received 02 January 2026 Revised 28 January 2026 Accepted 06 February 2026 Published 24 February 2026 Volume 17 - 2026 Edited by Yiju Wei, Shandong First Medical University, China Reviewed by Xingchen Peng, Sichuan University, China Lei Li, University of South China, China Updates Copyright 2026 Su, Yang and Zheng